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Nikon Z 400mm f/4.5 VR S Lens

A super-telephoto prime that increases your reach and features advanced optics, making it ideal for sport and wildlife applications. The Nikon Z 400mm f/4.5 VR S leaves you with the lightest lens in its class, two supported VR modes and uncompromised rendering capabilities. 

It offers outstanding Vibration Reduction (VR) with two optional VR modes - Normal and Sport, as well as Synchro VR when used with the Z9. To reduce blur while capturing handheld and achieve 5.5-stop increase in shutter speed, you can simply select the Normal mode. Professional photographers have praised the Sports mode for its ability to capture rapid and erratic moving subjects by tracking the subject and providing a stable viewfinder image throughout.

The S-Line optics delivers immense detail and groundbreaking anti-reflection performance to leave you with clear images even under harsh conditions. It also eliminates chromatic aberrations, colour fringing, and chromatic aberrations and provides one ED (extra-low dispersion) glass element, two Super ED elements and one SR (short-wavelength refractive) element. You can also increase the focal length as the existing 400mm can increase to 560mm or 800mm when a 1.4x or 2.0x teleconverter is attached.

  • Z-Mount Lens/FX Format
  • Aperture Range: f/4.5 to f/32
  • Super ED, ED, Fluorite, and SR Elements
  • Meso Amorphous and ARNEO Coatings
  • Silky Swift VCM Autofocus System
  • Vibration Reduction Image Stabilisation
  • Customisable Fn and Memory Set Buttons
  • Weather-Sealed Design

Experience quiet and accurate autofocus (AF) with a stepping motor (STM) that makes capturing wildlife and birds without disturbing them much easier.

Described as "the smallest and lightest 400mm Nikon has ever created," the NIKKOR Z 400mm f/4.5 VR S is a super-telephoto prime mixing unique portability with long reach, advanced optics, and apt AF and VR. It's a well-rounded lens for the travelling sports or wildlife photographer looking for a handheld design and capable reach for photographing distant subjects.

Long Reach, Compact Design

Distinct among super-telephoto lenses, this 400mm f/4.5 is especially sleek and lightweight, making it an ideal companion for travel or other photographing scenarios where handheld photographing is preferred. The sophisticated optical design includes a series of specialised elements that promote a lighter, more compact overall design. A modest f/4.5 maximum aperture offers a practical balance between size, speed, and weight. With a size more akin to a 70-200mm f/2.8, this lens measures just 234 mm long and weighs 1245 g. Compared to the Z 400mm f/2.8 lens, this f/4.5 version weighs 1723 g less and measures 152 mm shorter in length.

S-Line Optics

Helping to virtually eliminate chromatic aberrations, colour fringing, and chromatic aberrations, this lens features one ED (extra-low dispersion) glass element and two Super ED elements, along with one SR (short-wavelength refractive) element.

The SR element features a specialised high-dispersion design that refracts wavelengths shorter than blue; this helps achieve truly accurate colour rendering and also aids in making the lens design more compact.

Nano Crystal Coat has been applied for well-rounded anti-glare protection at all angles, helping to suppress ghosting, reflections, and flare for higher contrast and more accurate colour response when working in strong lighting and backlit conditions.

Autofocus and Vibration Reduction

Advanced autofocus system uses stepping motors to deliver quick and accurate focusing performance, along with smooth and quiet AF that is well-suited to both stills and video applications. Full-time manual focus override is also possible and an internal focusing design is used, where only the internal lens groups are moved during focusing, to maintain the overall length of the lens during use and to promote faster-focusing speeds.

Vibration Reduction image stabilisation reduces the appearance of camera shake by approximately 5.5 stops to better enable handheld shooting in low-light situations. Synchro VR is also supported when paired with compatible cameras featuring sensor-shift VR, which combines in-body VR with optical VR for improved results of up to 6 stops of compensation.

Professional Handling and Construction

As part of the lightweight, nimble design, the included tripod foot can easily be removed for a slimmed-down design that's ideal for handheld use. When used atop a tripod, the tripod mount has a rotating design for easy switching between horizontal and vertical orientations. An intuitive "no-look" control layout puts all of the frequently-used buttons and dials in easy-to-reach locations to prevent the need to move your eye from the viewfinder. Front anti-slip rubber grip for stable handheld use. A separate manual focus ring and a programmable control ring offer distinct levels of tactile control over photographing settings. Compatible with optional Z-Teleconverter TC-1.4x and TC-2x to further extend the reach of this lens.

The weather-sealed design includes rubber gaskets to prevent dust and moisture ingress and the exposed elements have a fluorine coating that repels dirt and oils and is easier to clean. Two customisable L-Fn buttons can be set to configure subject tracking, AF lock, image playback, or other modes. The L-Fn2 button is also accessible from four different locations on the barrel for easy reach. The focus limiter switch can constrain the focus range to 6 m to infinity or from the full focusing range of 2.5 m to infinity. A programmable Control Ring can also be used to control aperture, ISO, or exposure compensation. Kensington lock support for securing the lens in place during remote photographing sessions. Uses front screw-in 95mm filters.


Focal Length
Maximum Aperture
Minimum Aperture
Barrel Type
Camera Mount
Nikon Z
Full Time Manual Focus
Minimum Focus Distance
2.5 m
Internal Focus
Image Stabilisation
Yes, Vibration Reduction
Filter Thread
95 mm (Front)
1245 g
R 75,995 Only 1 left in Store
Payment plans available
Delivery Truck Free Standard shipping included. Faster options available

What's in the box

  • Nikon NIKKOR Z 400mm f/4.5 VR S Lens
    • Nikon LC-95B 95mm Snap-On Front Lens Cap
    • Nikon LF-N1 Rear Lens Cap
    • Nikon HB-105 Lens Hood
    • Nikon CL-C5 Lens Case
    • Tripod Foot
  • Limited 2-Year Warranty

Reviewed by 0 customers

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